The TV campaign, which breaks today and runs until July 31, will feature Kevin Ralph, the manager of its Archway, London, branch, explaining key messages to counter allegations that the food is unhealthy.
This is the first time the fast-food giant has used an employee rather than an actor in its advertising, but a spokeswoman added that, "it was too soon to say" whether Kevin would have the same impact as Halifax's Howard, who has gone on to become a minor personality releasing a record and giving press interviews.
Kevin was selected from a raft of staff recommendations made by area managers across the UK.
This latest push is part of an ongoing customer consultation campaign, launched by chairman and CEO Peter Beresford last year.
Also launched today is a print campaign in papers including The Times, The Sun and the Daily Mirror featuring a full page ad, with a message from Beresford saying: "Like any restaurant we have our critics. Like any good restaurant we listen".
The message continues that following feedback from customers changes have now been made such as children being offered carrots with their meals.
A spokeswoman said: "This is about explaining to people some of the things they didn't know, such as our use of local producers."
The TV and print campaigns feature the slogan "we hear you".
The spokeswoman added that the print ad may also run in some Sunday papers this weekend, as well as selected women's titles.
Among the fast-food chain's most prominent critics is film-maker Morgan Spurlock, whose film 'Super Size Me', in which he lived on a McDonald's diet for a month, was a hit with UK audiences last year.
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