It fell from 358,345 copies in July to 341,698 copies in August. On Monday, it is changing to its new size, halfway between a broadsheet and a tabloid, which it hopes will bring the circulation boost enjoyed by the already-switched Times and Independent.
The surprise for those two was a 2.55% fall in The Times' circulation, to 680,214, reversing its upward trend. The Independent kept its head above water with a 0.12% increase to 255,906.
Of the remaining broadsheets, The Daily Telegraph experienced a 0.84% fall to 904,660 and the Financial Times a 0.87% rise to 413,882.
In the mid-market, the Daily Mail fell 1.47% to 2,384,943 while the Daily Express rose by an impressive 4.86% to 876,563.
London's Evening Standard lost 15.15% of its circulation in a much quieter news month compared with the aftermath of the July 7 terrorist attacks. It sold 316,453 copies in August, compared with 332,524 in June and 372,955 in July. The news comes amid speculation that it is to ditch its freesheet version of the newspaper, Standard Lite.
It was a good month for the tabloids in that none saw circulation falls, although the rises were nothing to shout about.
The Sun was up 0.53% to 3,361,306; the Mirror was up 0.96% to 1,769,771; and the Daily Star was up 0.42% to 893,601.
In the Sunday market, Guardian Newspapers' pain was doubled by The Observer's 4.49% fall to 425,737.
The Sunday Times eclipsed its daily stablemate with a 1.44% rise to 1,357,916 copies, while The Independent on Sunday sagged to 202,248, a fall of 2.15%. The Sunday Telegraph was down 1.93% to 669,747.
The Sunday Express led the mid-market with a 3.63% rise to 919,583, while the Daily Mail was up just 0.45% to 2,272,476.
The Sunday Mirror was up 0.61% to 1,558,367 while the News of the World was up 1.55% to 3,759,443. The People was the only faller, down 3.11% to 945,803.
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