The TV, print, outdoor and online campaign, which breaks next Monday, is designed to inform people that the animals packages are now available on prepay as well as pay monthly.
The previously used motif of balloons representing the four different packages recurs in the TV spot.
The ad shows a crowded urban train station as two trains arrive. When the doors open, thousands of the animal-shaped balloons float out, and the people start to pick out the balloon they like best.
Tony McTear, creative director at Fallon London, said: "We set out to give the Animals a fresh spin and energy that signals something new is happening at Orange.
"The story could've been easy to over complicate, but we wanted to keep the idea simple and especially fun."
Orange introduced a new animal to its pay-as-you-go line-up, Camel, which is designed for customers with friends and family abroad. The other three options are Dolphin, Raccoon and Canary, which also feature on pay monthly with Panther.
Justin Billingsley, director of brand marketing at Orange UK, said: "This campaign highlights how everybody is different and that our animal packages let our pay-as-you-go customers choose the reward that best suits them."
Orange has 10m pay-as-you-go customers in the UK.
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