Despite the fall, The Guardian remains just above the 400,000 mark according to November's ABC figures.
After changing format on September 12, The Guardian hit a high of 404,187 copies during September as readers sampled the new look, but after a small fall in October it fell again by 0.56% in November to 401,029 copies.
The Times, which has enjoyed a sustained boost from its conversion to compact in November 2004, fell 1.74% from 703,792 to 691,283. The Independent also lost circulation, shifting 1.34% less than in October at 263,449.
Broadsheet perseverer The Daily Telegraph held its own with a 0.19% rise to 903,405, shrugging off editor Martin Newland's sudden departure in the middle of the month. The paper is now looking for a new editor.
The Financial Times, which also parted company with its editor Andrew Gowers in November, rose 3% to 431,806.
The Daily Mail slipped 0.39% to 2.341m, but the Daily Express fared worse, going under 800,000 with a 1.76% drop to 796,592.
The red-top sector also declined, with The Sun down 0.98% to 3.193m, the Daily Mirror down 0.75% to 1.672m, and the Daily Star down 3.01% to 795,308.
Among the dailies, the London Evening Standard had the best month, rising 5.24% to 346,489.
In the Sunday market, editor Sarah Sands' makeover of the Sunday Telegraph brought an 8.1% rise in circulation, shifting around 53,000 extra copies to reach 714,992. The Sunday Times absorbed the impact fairly well, losing just under 10,000 copies, or 0.68%, to hit 1,395,046.
However, Observer readers may have been lured away -- it sank by just under 15,000 copies, or 3.3%, to 435,882 as its plans for a Berliner makeover draw nearer.
The Independent on Sunday did not sustain the rise it enjoyed last month with its conversion to the tabloid format. It lost more than 4,000 of the 27,000 copies it added, leaving it 1.92% down at 225,629.
The Sunday Express put on 5.1% to reach 871,312 and The Mail on Sunday put on 1.98% to reach 2.338m.
The red-tops were the only category to lose overall circulation, with the News of the World down 1.08% to 3.733m, the Sunday Mirror down 2.5% to 1.421m, and The People down 3.88% to 870,389.
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