The Daily Express was down the most in percentage terms, by 7% to 1,742,000.
The Times also dropped 7% to 1,730,000, while other quality press showed gains.
The Financial Times underlined its recent recovery with a 13% rise to 394,000. The Daily Telegraph, The Guardian and The Independent all rose 5%.
However, the Daily Telegraph was overtaken as the UK's fifth most-read newspaper by Metro, which gained 16% to 2,195,000. The Daily Telegraph has a readership of 2,177,000.
The most significant changes among the Sunday papers were a 9% rise for The Observer to 1,445,000 and an 11% drop for The People to 1,756,000.
The figures for magazines illustrated the readership trend away from classic women's weeklies to newer titles.
There were large jumps for ACP-Natmag's 2004 launch Reveal, up 84% to 717,000; Emap's 2002 launch Closer, was up 21% to 983,000; and IPC's January 2005 launch Pick Me Up was up 21% to 983,000.
IPC's Woman's Own, launched in 1932, was down 21% to 1,213,000, and its title Woman, launched in 1937, was down 21% to 924,000.
The figures show men's magazines declining, but perhaps not as much as observers feared. The market leader, Emap's FHM, lost 8% of its readership, falling to 2,397,000, while IPC's Loaded dropped 6% to 1,061,000 and Dennis' Maxim dropped 9% to 602,000.