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Zoo's sales rose by 30.1% year on year to 260,317, but it still trails arch-rival Nuts, which increased sales by a more modest 5% to 304, 751.

Highbury House's Front recorded an increase of 4.8% for the period, but its year-on-year sales dropped by 12.1% to 88,154.

MediaCom director of press Steve Goodman said this fall could be due to readers moving to Nuts and Zoo, as Front's editorial content is more similar in tone to the weeklies than that of any other men's monthly title.

Sales of the market leader, Emap's FHM, were down by 2.4% year on year to 560,167.

NatMags' upmarket title, Esquire, recorded a more notable drop in circulation of 10.9% in the period, following a cover-price hike.

The original lads' mag, Loaded, regained its position as the number two monthly, with a 7.7% sales rise, after a spring relaunch and price cut.

In the previous ABC period, the title had slipped to fourth place behind Dennis Publishing's Maxim and Natmag Rodale's Men's Health.

Bruce Sandell, publishing director of Maxim, said the magazine would be running an ad campaign, through Flint Creative, in the autumn.

FHM Worldwide managing director David Pullan said the ABCs showed weeklies and monthlies could 'co-exist peacefully'. 'It is clear that the weeklies are bringing more men into buying magazines,' he added.

Haymarket's Stuff grew by 10% year on year to 77,373, a ninth-consecutive sales rise.